Sphenodon
Sphenodon is the only living representative of the order
Rhynchocephalia, of class Reptilia. It was once widely distributed in the main
islands of New Zealand but is now restricted to some small island in the Bay of
Plenty, North Islands. As it shows many primitive features and shares some
amphibian and reptilian characters, it is often called the ‘living fossil’
forming a sort of bridge between the two groups of vertebrates. Sphenodon is
called ‘Tuatara’ by the natives. Sphenodon is a rather large lizard like
reptile. It is nocturnal in habit and lives in burrows in the daytime. It feeds
mainly on insects. Dorsally the body is dull olive green with yellow spots and
ventrally it is whitish. Small granular scales cover the body, while on the
ventral side there are transverse rows of large square plates formed by scales.
A mid-dorsal row of scales forms spines (frill) from the head to the end of the
tail, except on the neck. The tail is thick and laterally compressed. The eyes
are large with vertical pupil. The males lack copulatory organs. The cloacal
aperture is in the form of a transverse slit.
The skull of sphenodon consists of paired premaxillae, frontals,
nasals and parietals. Three temporal fossae are present. The quadrate is fixed.
The pterygoids extend forwards separating the palatines. Post-frontals are
separate. Teeth are acrodont and are found in two parallel rows. The vertebrae
are amphicoelous with persistent notochord. A pro-atlas is present between the
skull and atlas bone. Ribs are single-headed and abdominal ribs are present. A median
sternum is present. Caudal vertebrae have chevron bones. In the forelimbs
radius and ulna are separate. A median interclavicles and a ventro-median
procoracoid are present. A cartilaginous epipubis is present. The ilia are
blade- like. The heart is of reptilian type. But the three principal arches
arise from a short common trunk. The arterial and venous systems show
similarities with those of Urodela. The brain has well-developed central
hemispheres and optic lobes, but the cerebellum is poorly developed. There is a
distinct ‘parietal eye’ with lens, retina and nerve connection with brain. In the
adult the parietal foramen is covered with skin. It is the only reptile in
which the male lacks penis. The eggs take about 13 months to hatch.
The anatomical features of sphenodon clearly indicate that it is
a reptile, though primitive and almost ideally generalized. But its true systematic
position has been a matter of controversy and opinions differ among the
different zoologists.
Reference
Also See
• Parrots
• Bats
• Bears
• Birds
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